View source for Nutrition

{{Short description|Provision to cells and organisms to support life}}
{{Other uses}}
{{Use American English|date=September 2025}}
{{Use dmy dates|date=August 2022}}
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[[File:Close wing posture of Amblypodia anita (Hewitson, 1862) - Purple Leaf Blue (Male) gathering nutrients from bird dropping WLB.jpg|thumb|upright=1.5|alt=see caption|A purple leaf blue butterfly (''[[Amblypodia anita]]'') gathering [[nutrient]]s from [[guano]]]]

'''Nutrition''' is the [[biochemistry|biochemical]] and [[physiology|physiological]] process by which an [[organism]] uses [[food]] and [[water]] to support its [[life]]. The intake of these substances provides organisms with [[nutrient]]s (divided into [[Macronutrient|macro-]] and [[Micronutrient|micro-]]) which can be [[Metabolism|metabolized]] to create [[Food energy|energy]] and chemical structures; too much or too little of an essential nutrient can cause [[malnutrition]]. [[Nutritional science]], the study of nutrition as a [[hard science]], typically emphasizes [[human nutrition]].

The type of organism determines what nutrients it needs and how it obtains them. Organisms obtain nutrients by consuming [[organic matter]], consuming inorganic matter, absorbing light, or some combination of these. Some can produce nutrients internally by consuming basic elements, while others must consume other organisms to obtain pre-existing nutrients. All forms of life require [[carbon]], [[Biological thermodynamics|energy]], and [[water]] as well as various other [[molecule]]s. Animals require complex nutrients such as [[carbohydrate]]s, [[lipid]]s, and [[protein]]s, obtaining them by consuming other organisms. Humans have developed [[agriculture]] and [[cooking]] to replace [[foraging]] and advance human nutrition. Plants acquire nutrients through the [[soil]] and the atmosphere. Fungi absorb nutrients around them by breaking them down and absorbing them through the [[mycelium]].

== History ==
{{Main|Nutritional science}}
Scientific analysis of food and nutrients began during the [[chemical revolution]] in the late 18th century. Chemists in the 18th and 19th centuries experimented with different elements and food sources to develop theories of nutrition.<ref>{{Cite journal |last=Carpenter |first=Kenneth J. |date=1 March 2003 |title=A Short History of Nutritional Science: Part 1 (1785–1885) |journal=The Journal of Nutrition |volume=133 |issue=3 |pages=638–645 |doi=10.1093/jn/133.3.638 |pmid=12612130 |doi-access=free }}</ref> Modern nutrition science began in the 1910s as individual micronutrients began to be identified. The first vitamin to be chemically identified was [[thiamine]] in 1926, and vitamin C was identified as a protection against scurvy in 1932.<ref>{{Cite journal |last=Stafford |first=Ned |date=December 2010 |title=History: The changing notion of food |journal=Nature |volume=468 |issue=7327 |pages=S16–S17 |doi=10.1038/468S16a |pmid=21179078 |doi-access=free |bibcode=2010Natur.468S..16S }}</ref> The role of vitamins in nutrition was studied in the following decades. The first recommended dietary allowances for humans were developed to address fears of disease caused by food deficiencies during the [[Great Depression]] and the Second World War.<ref>{{Cite journal |last1=Mozaffarian |first1=Dariush |last2=Rosenberg |first2=Irwin |last3=Uauy |first3=Ricardo |date=13 June 2018 |title=History of modern nutrition science—implications for current research, dietary guidelines, and food policy |journal=[[BMJ]] |volume=361 |article-number=k2392 |doi=10.1136/bmj.k2392 |pmc=5998735 |pmid=29899124 }}</ref> Due to its importance in human health, the study of nutrition has heavily emphasized human nutrition and agriculture, while [[ecology]] is a secondary concern.{{Sfn|Simpson|Raubenheimer|2012|p=2}}

== Nutrients ==
{{main|Nutrient}}

{{multiple image
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| footer            = [[Composting]] within [[agricultural systems]] capitalizes upon the natural services of nutrient recycling in ecosystems. [[Bacteria]], [[fungi]], [[insects]], [[earthworms]], [[Hemiptera|bugs]], and other creatures dig and digest the compost into fertile soil. The minerals and nutrients in the soil are recycled back into the production of crops.
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Nutrients are substances that provide energy and physical components to the organism, allowing it to survive, grow, and reproduce. Nutrients can be basic elements or complex [[macromolecule]]s. Approximately 30 elements are found in [[organic matter]], with [[nitrogen]], [[carbon]], and [[phosphorus]] being the most important.{{Sfn|Andrews|2017|pp=70–72}} [[Macronutrient]]s are the primary substances required by an organism, and [[micronutrient]]s are substances required by an organism in trace amounts. Organic micronutrients are classified as [[vitamin]]s, and inorganic micronutrients are classified as [[mineral (nutrition)|minerals]]. Over-nutrition of macronutrients is a major cause of obesity and increases the risk of developing various non-communicable diseases (NCDs), including type 2 diabetes, stroke, hypertension, coronary heart disease, osteoporosis, and some forms of cancer.<ref>{{Cite book |last=Gibney |first=Michael J. |url=https://www.google.co.uk/books/edition/Introduction_to_Human_Nutrition/Pn7J9JbE4g0C?hl=en&gbpv=0 |title=Introduction to Human Nutrition |last2=Lanham-New |first2=Susan A. |last3=Cassidy |first3=Aedin |last4=Vorster |first4=Hester H. |date=2013-03-14 |publisher=John Wiley & Sons |isbn=978-1-118-68470-2 |pages=350 |language=en}}</ref> Nutrients can also be classified as essential or nonessential, with essential meaning the body cannot synthesize the nutrient on its own.{{Sfn|Wu|2017|pp=2–4}}

Nutrients are absorbed by the cells and used in metabolic biochemical reactions. These include fueling reactions that create precursor [[metabolite]]s and energy, biosynthetic reactions that convert precursor metabolites into building block molecules, [[polymerization]]s that combine these molecules into macromolecule [[polymer]]s, and assembly reactions that use these polymers to construct cellular structures.{{Sfn|Andrews|2017|pp=70–72}}

=== Nutritional groups ===
{{main|Primary nutritional groups}}
Organisms can be classified by how they obtain carbon and energy. [[Heterotroph]]s are organisms that obtain nutrients by consuming the carbon of other organisms, while [[autotroph]]s are organisms that produce their own nutrients from the carbon of inorganic substances like [[carbon dioxide]]. [[Mixotroph]]s are organisms that can be heterotrophs and autotrophs, including some [[plankton]] and [[carnivorous plant]]s. [[Phototroph]]s obtain energy from light, while [[chemotroph]]s obtain energy by consuming chemical energy from matter. [[Organotroph]]s consume other organisms to obtain electrons, while [[lithotroph]]s obtain electrons from inorganic substances, such as [[water]], [[hydrogen sulfide]], [[dihydrogen]], [[iron(II)]], [[sulfur]], or [[ammonium]].{{Sfn|Andrews|2017|pp=72–79}} Prototrophs can create essential nutrients from other compounds, while [[auxotroph]]s must consume preexisting nutrients.{{Sfn|Andrews|2017|p=93}}

=== Diet ===
{{main|Diet (nutrition)}}In nutrition, the [[Diet (nutrition)|diet]] of an organism is the sum of the foods it eats.<ref>{{Cite web |title=Diet |url=https://education.nationalgeographic.org/resource/diet |access-date=8 August 2022 |website=[[National Geographic]]}}</ref> A healthy diet improves the physical and mental health of an organism. This requires ingestion and absorption of [[vitamin]]s, [[Dietary mineral|minerals]], [[essential amino acid]]s from protein and [[essential fatty acid]]s from fat-containing food. [[Carbohydrates]], protein and fat play major roles in ensuring the [[quality of life]], [[health]] and [[longevity]] of the organism.<ref>{{Cite web |date=2021-05-16 |title=Benefits of Healthy Eating |url=https://www.cdc.gov/nutrition/resources-publications/benefits-of-healthy-eating.html |access-date=2023-05-17 |website=Centers for Disease Control and Prevention |language=en-us}}</ref> Some cultures and religions have restrictions on what is acceptable for their diet.<ref>{{Cite web |date=2016-09-19 |title=Religion and dietary choices |url=https://www.independentnurse.co.uk/content/clinical/religion-and-dietary-choices/ |access-date=2023-05-17 |website=Independent Nurse |language=en}}</ref>

=== Nutrient cycle ===
{{main|Nutrient cycle}}A [[nutrient cycle]] is a [[biogeochemical cycle]] involving the movement of inorganic matter through a combination of soil, organisms, air or water, where they are exchanged in organic matter.<ref name=":0">{{cite encyclopedia |last=Allaby |first=Michael |year=2015 |encyclopedia=A Dictionary of Ecology |title=Nutrient cycle |publisher=[[Oxford University Press]] |location=Oxford |edition=5th |url=https://www.oxfordreference.com/view/10.1093/acref/9780191793158.001.0001/acref-9780191793158-e-3868 |isbn=978-0-19-179315-8 |url-access=subscription}} {{subscription required}}</ref> Energy flow is a unidirectional and noncyclic pathway, whereas the movement of [[mineral nutrients]] is cyclic. Mineral cycles include the [[carbon cycle]], [[sulfur cycle]], [[nitrogen cycle]], [[water cycle]], [[phosphorus cycle]], and [[oxygen cycle]], among others that continually recycle along with other mineral nutrients into [[Productivity (ecology)|productive]] ecological nutrition.<ref name=":0" />

[[Biogeochemical cycle]]s that are performed by living organisms and natural processes are [[Water cycle|water]], [[Carbon cycle|carbon]], [[Nitrogen cycle|nitrogen]], [[Phosphorus cycle|phosphorus]], and [[sulfur cycle]]s.<ref>{{Cite web |title=Intro to biogeochemical cycles (article) |url=https://www.khanacademy.org/science/biology/ecology/biogeochemical-cycles/a/introduction-to-biogeochemical-cycles |access-date=2022-11-02 |website=Khan Academy |language=en}}</ref> Nutrient cycles allow these essential elements to return to the environment after being absorbed or consumed.<ref>{{cite web | url=https://bio.libretexts.org/Bookshelves/Botany/Botany_(Ha_Morrow_and_Algiers)/Unit_3%3A_Plant_Physiology_and_Regulation/15%3A_Nutrition_and_Soils/15.03%3A_Nutrient_Cycles#:~:text=Nutrient%20cycles%2C%20also%20known%20as,to%20plants%20and%20other%20organisms | title=4.3.3: Nutrient Cycles | date=29 May 2020 }}</ref> Without proper nutrient cycling, there would be a risk of change in oxygen levels, climate, and ecosystem function.{{fact|date=November 2024}}

=== Foraging ===
{{Main|Foraging}}
[[File:BonoboFishing04.jpeg|thumb|A [[bonobo]] fishing for [[termite]]s with a prepared stick]]
Foraging is the process of seeking out nutrients in the environment. It may also be defined to include the subsequent use of the resources. Some organisms, such as animals and bacteria, can navigate to find nutrients, while others, such as plants and fungi, extend outward to find nutrients. Foraging may be random, in which the organism seeks nutrients without method, or it may be systematic, in which the organism can go directly to a food source.{{Sfn|Andrews|2017|pp=83–85}} Organisms are able to detect nutrients through [[taste]] or other forms of [[nutrient sensing]], allowing them to regulate nutrient intake.{{Sfn|Simpson|Raubenheimer|2012|p=36}} [[Optimal foraging theory]] is a model that explains foraging behavior as a [[cost–benefit analysis]] in which an animal must maximize the gain of nutrients while minimizing the amount of time and energy spent foraging. It was created to analyze the foraging habits of animals, but it can also be extended to other organisms.{{Sfn|Andrews|2017|p=16}} Some organisms are specialists that are adapted to forage for a single food source, while others are generalists that can consume a variety of food sources.{{Sfn|Andrews|2017|p=98}}

== Nutrient deficiency ==
{{See also|Malnutrition}}
Nutrient deficiencies, known as [[malnutrition]], occur when an organism does not have the nutrients that it needs. A deficiency is not the same as a nutrient inadequacy which occurs when the intake of nutrients is above the level of deficiency, but below the recommended dietary level. This may lead to hidden symptoms of nutrient deficiency that are difficult to identify.<ref>{{Cite journal |last=Kiani |first=Aysha Karim |last2=Dhuli |first2=Kristjana |last3=Donato |first3=Kevin |last4=Aquilanti |first4=Barbara |last5=Velluti |first5=Valeria |last6=Matera |first6=Giuseppina |last7=Iaconelli |first7=Amerigo |last8=Connelly |first8=Stephen Thaddeus |last9=Bellinato |first9=Francesco |last10=Gisondi |first10=Paolo |last11=Bertelli |first11=Matteo |date=2022 |title=Main nutritional deficiencies |journal=Journal of Preventive Medicine and Hygiene |volume=63 |issue=2 Suppl 3 |article-number=93 |doi=10.15167/2421-4248/jpmh2022.63.2S3.2752 |issn=2421-4248 |pmc=9710417 |pmid=36479498}}</ref> Nutrient deficiency may be caused by a sudden decrease in nutrient intake or by an inability to absorb essential nutrients. Not only is malnutrition the result of a lack of necessary nutrients,<ref>{{cite journal |last1=Mora |first1=Rafael J.F. |title=Malnutrition: Organic and Functional Consequences |journal=World Journal of Surgery |date=June 1999 |volume=23 |issue=6 |pages=530–535 |doi=10.1007/PL00012343 |pmid=10227920 }}</ref> but it can also be a result of other illnesses and health conditions. When this occurs, an organism will adapt by reducing energy consumption and expenditure to prolong the use of stored nutrients. It will use stored energy reserves until they are depleted.<ref>{{Cite journal |last=Mora |first=Rafael J.F. |date=1 June 1999 |title=Malnutrition: Organic and Functional Consequences |journal=[[World Journal of Surgery]] |volume=23 |issue=6 |pages=530–535 |doi=10.1007/PL00012343 |pmid=10227920 }}</ref>

A balanced diet includes appropriate amounts of all essential and non-essential nutrients. These can vary by age, weight, sex, physical activity levels, and more. A lack of just one essential nutrient can cause bodily harm, just as an overabundance can cause toxicity. The Daily Reference Values keep the majority of people from nutrient deficiencies.<ref>{{cite web | url=https://www.efsa.europa.eu/en/topics/topic/dietary-reference-values#:~:text=DRVs%20are%20key%20concepts%20in,at%20population%20and%20individual%20level | title=Dietary reference values &#124; EFSA | date=5 August 2024 }}</ref> DRVs are not recommendations but a combination of nutrient references to educate professionals and policymakers on what the maximum and minimum nutrient intakes are for the average person.<ref>{{cite web | url=https://www.efsa.europa.eu/en/topics/topic/dietary-reference-values | title=Dietary reference values &#124; EFSA | date=5 August 2024 }}</ref> Food labels also use DRVs as a reference to create safe nutritional guidelines for the average healthy person.<ref>{{Cite journal |last=Nutrition |first=Center for Food Safety and Applied |date=2024-03-13 |title=Daily Value on the Nutrition and Supplement Facts Labels |url=https://www.fda.gov/food/nutrition-facts-label/daily-value-nutrition-and-supplement-facts-labels |archive-url=https://web.archive.org/web/20231001072833/https://www.fda.gov/food/nutrition-facts-label/daily-value-nutrition-and-supplement-facts-labels |url-status=dead |archive-date=1 October 2023 |journal=FDA |language=en}}</ref>

== In organisms ==

=== Animal ===
{{main|Animal nutrition}}
[[File:Alcedo atthis eating a tadpole.jpg|alt=see caption|thumb|A [[kingfisher]] eating a [[tadpole]] near the [[Ariège (river)|Ariège river]], France]]
Animals are heterotrophs that consume other organisms to obtain nutrients. [[Herbivore]]s are animals that eat plants, [[carnivore]]s are animals that eat other animals, and [[omnivore]]s are animals that eat both plants and other animals.{{Sfn|Wu|2017|p=1}} Many herbivores rely on bacterial fermentation to create digestible nutrients from indigestible plant cellulose, while obligate carnivores must eat animal meats to obtain certain vitamins or nutrients their bodies cannot otherwise synthesize. Animals generally have a higher requirement of energy in comparison to plants.<ref>{{cite web |last1=National Geographic Society |date=21 January 2011 |title=Herbivore |url=https://www.nationalgeographic.org/encyclopedia/herbivore/ |access-date=1 May 2017 |website=[[National Geographic Society]] |language=en |archive-date=25 February 2017 |archive-url=https://web.archive.org/web/20170225044420/http://www.nationalgeographic.org/encyclopedia/herbivore/ |url-status=live|author1-link=National Geographic Society }}</ref> The macronutrients essential to animal life are carbohydrates, [[amino acid]]s, and [[fatty acid]]s.{{Sfn|Wu|2017|pp=2–4}}<ref>{{Cite web |title=Nutrition: What Plants and Animals Need to Survive {{!}} Organismal Biology |url=https://organismalbio.biosci.gatech.edu/nutrition-transport-and-homeostasis/nutrition-needs-and-adaptations/ |access-date=2022-11-02 |website=organismalbio.biosci.gatech.edu}}</ref>

All macronutrients except water are required by the body for energy, however, this is not their sole physiological function. The energy provided by macronutrients in food is measured in kilocalories, usually called Calories, where 1 Calorie is the amount of energy required to raise 1 kilogram of water by 1 degree Celsius.<ref>{{Cite web |date=2018-12-21 |title=4.2: Nutrients |url=https://bio.libretexts.org/Bookshelves/Human_Biology/Book%3A_Human_Biology_(Wakim_and_Grewal)/04%3A_Nutrition/4.2%3A_Nutrients |access-date=2022-11-02 |website=Biology LibreTexts |language=en}}</ref>

Carbohydrates are molecules that store significant amounts of energy. Animals digest and metabolize carbohydrates to obtain this energy. Carbohydrates are typically synthesized by plants during metabolism, and animals have to obtain most carbohydrates from nature, as they have only a limited ability to generate them. They include [[sugar]]s, [[oligosaccharide]]s, and [[polysaccharide]]s. [[Glucose]] is the simplest form of carbohydrate.{{Sfn|Mann|Truswell|2012|pp=21–26}} Carbohydrates are broken down to produce glucose and [[short-chain fatty acid]]s, and they are the most abundant nutrients for herbivorous land animals.{{Sfn|Wu|2017|pp=193–194}} Carbohydrates contain 4 calories per gram.

Lipids provide animals with fats and oils. They are not soluble in water, and they can store energy for an extended period of time. They can be obtained from many different plant and animal sources. Most dietary lipids are [[triglyceride]]s, composed of [[glycerol]] and fatty acids. [[Phospholipid]]s and [[sterol]]s are found in smaller amounts.{{Sfn|Mann|Truswell|2012|pp=49–55}} An animal's body will reduce the amount of fatty acids it produces as dietary fat intake increases, while it increases the amount of fatty acids it produces as carbohydrate intake increases.{{Sfn|Wu|2017|p=271}} Fats contain 9 calories per gram.

Protein consumed by animals is broken down to amino acids, which would be later used to synthesize new proteins. Protein is used to form cellular structures, fluids,{{Sfn|Mann|Truswell|2012|pp=70–73}} and [[enzyme]]s (biological [[catalyst]]s). Enzymes are essential to most [[Metabolism|metabolic]] processes, as well as [[DNA replication]], [[DNA repair|repair]], and [[Transcription (genetics)|transcription]].<ref name="EXPASY2000">{{cite journal |vauthors=Bairoch A |date=January 2000 |title=The ENZYME database in 2000 |journal=[[Nucleic Acids Research]] |volume=28 |issue=1 |pages=304–05 |doi=10.1093/nar/28.1.304 |pmc=102465 |pmid=10592255 }}</ref> Protein contains 4 calories per gram.

Much of [[Ethology|animal behavior]] is governed by nutrition. [[Animal migration|Migration patterns]] and [[Seasonal breeder|seasonal breeding]] take place in conjunction with food availability, and [[courtship display]]s are used to display an animal's health.{{Sfn|Simpson|Raubenheimer|2012|pp=3–4}} Animals develop positive and negative associations with foods that affect their health, and they can instinctively avoid foods that have caused [[toxic injury]] or nutritional imbalances through a [[conditioned food aversion]]. Some animals, such as rats, do not seek out new types of foods unless they have a nutrient deficiency.{{Sfn|Simpson|Raubenheimer|2012|pp=39–41}}

==== Human ====
{{main|Human nutrition|Cooking}}
Early human nutrition consisted of foraging for nutrients, like other animals, but it diverged at the beginning of the [[Holocene]] with the [[Neolithic Revolution]], in which humans developed agriculture to produce food. The Chemical Revolution in the 18th century allowed humans to study the nutrients in foods and develop more advanced methods of [[food preparation]]. Major advances in economics and technology during the 20th century allowed [[mass production]] and [[food fortification]] to better meet the nutritional needs of humans.<ref>{{Cite book |last=Trüeb |first=Ralph M. |title=Nutrition for Healthy Hair |publisher=[[Springer Science+Business Media|Springer]] |year=2020 |isbn=978-3-030-59920-1 |pages=3–15 |chapter=Brief History of Human Nutrition |doi=10.1007/978-3-030-59920-1_2 }}</ref> [[Human behavior]] is closely related to human nutrition, making it a subject of [[social science]] in addition to biology. Nutrition in humans is balanced with eating for pleasure, and optimal diet may vary depending on the demographics and health concerns of each person.{{Sfn|Mann|Truswell|p=1|2012}} Social determinants of health (SDOH) and structural factors drive nutrition and diet-related health disparities.<ref>{{Cite journal |last=Kahlon |first=Maninder |date=2024-04-01 |title=Understanding and Intervening in Nutrition-Related Health Disparities |url=https://www.sciencedirect.com/science/article/pii/S2161831324000292 |journal=Advances in Nutrition |volume=15 |issue=4 |pages=2 |doi=10.1016/j.advnut.2024.100195 |issn=2161-8313|pmc=11031373 }}</ref> 

Humans are omnivores that eat a variety of foods. Cultivation of [[cereal]]s and production of [[bread]] has made up a key component of human nutrition since the beginning of agriculture. Early humans hunted animals for meat, and modern humans domesticate animals to consume their meat and eggs. The development of [[animal husbandry]] has also allowed humans in some cultures to consume the [[milk]] of other animals and process it into foods such as [[cheese]]. Other foods eaten by humans include nuts, seeds, fruits, and vegetables. Access to domesticated animals as well as [[vegetable oil]]s has caused a significant increase in human intake of fats and oils. Humans have developed advanced methods of [[food processing]] that prevent contamination of pathogenic microorganisms and simplify the production of food. These include drying, freezing, heating, milling, pressing, packaging, refrigeration, and irradiation. Most cultures add [[herb]]s and [[spice]]s to foods before eating to add flavor, though most do not significantly affect nutrition. Other additives are also used to improve the safety, quality, flavor, and nutritional content of food.{{Sfn|Mann|Truswell|pp=409–437|2012}}

Humans obtain most carbohydrates as starch from cereals, though sugar has grown in importance.{{Sfn|Mann|Truswell|2012|pp=21–26}} Lipids can be found in [[animal fat]], [[butterfat]], vegetable oil, and [[leaf vegetable]]s, and they are also used to increase flavor in foods.{{Sfn|Mann|Truswell|2012|pp=49–55}} Protein can be found in virtually all foods, as it makes up cellular material, though certain methods of food processing may reduce the amount of protein in a food.{{Sfn|Mann|Truswell|2012|p=86}} Humans can also obtain energy from [[ethanol]], which is both a food and a drug, but it provides relatively few essential nutrients and is associated with nutritional deficiencies and other health risks.{{Sfn|Mann|Truswell|2012|pp=109–120}}

In humans, poor nutrition can cause deficiency-related diseases, such as [[blindness]], [[anemia]], [[scurvy]], [[preterm birth]], [[stillbirth]] and [[cretinism]],<ref name="Whitney-Rolfes-lead">{{cite book |last1=Whitney |first1=Ellie |title=Understanding Nutrition |last2=Rolfes |first2=Sharon Rady |publisher=Wadsworth, [[Cengage Learning]] |year=2013 |isbn=978-1-133-58752-1 |edition=13 |pages=667, 670}}</ref> or nutrient-excess conditions, such as [[obesity]]<ref>{{cite web |date=7 June 2021 |title=Defining Adult Overweight and Obesity |url=https://www.cdc.gov/obesity/adult/defining.html |website=[[Centers for Disease Control and Prevention]] |access-date=22 September 2021 |archive-date=28 September 2021 |archive-url=https://web.archive.org/web/20210928174006/https://www.cdc.gov/obesity/adult/defining.html |url-status=live}}</ref> and [[metabolic syndrome]].<ref>{{cite web |title=Metabolic syndrome – PubMed Health |publisher=[[National Center for Biotechnology Information]] |url=http://www.ncbi.nlm.nih.gov/pubmedhealth/PMH0004546 |archive-url=https://web.archive.org/web/20110205042451/http://www.ncbi.nlm.nih.gov/pubmedhealth/PMH0004546 |archive-date=5 February 2011 |url-status=dead |access-date=17 October 2011}}</ref> Other conditions possibly affected by nutrition disorders include [[cardiovascular disease]]s,<ref>{{cite web|url=http://www.umm.edu/altmed/articles/omega-3-000316.htm |title=Omega-3 fatty acids |archive-url=https://web.archive.org/web/20080709084934/http://www.umm.edu/altmed/articles/omega-3-000316.htm |archive-date=9 July 2008 |website=Umm.edu |date=5 October 2011 |access-date=17 October 2011}}</ref> [[diabetes]],<ref>{{cite book |url=https://permanent.fdlp.gov/lps125227/Eating_Diabetes.pdf |title=What I need to know about eating and diabetes |date=2007 |publisher=U.S. Department of Health and Human Services, National Institutes of Health, National Institute of Diabetes and Digestive and Kidney Diseases, National Diabetes Information Clearinghouse |oclc=656826809 |access-date=22 September 2021 |archive-date=8 August 2022 |archive-url=https://web.archive.org/web/20220808135741/https://permanent.fdlp.gov/lps125227/Eating_Diabetes.pdf |url-status=live}}</ref><ref>{{cite web|url=http://www.helpguide.org/life/healthy_diet_diabetes.htm |title=Diabetes Diet and Food Tips: Eating to Prevent and Control Diabetes |archive-url=https://web.archive.org/web/20110520045538/http://www.helpguide.org/life/healthy_diet_diabetes.htm |archive-date=20 May 2011 |website=Helpguide.org |access-date=17 October 2011}}</ref> and [[osteoporosis]].<ref>{{cite web|url=https://ods.od.nih.gov/factsheets/VitaminD-HealthProfessional/ |title=Dietary Supplement Fact Sheet: Vitamin D |archive-url=https://web.archive.org/web/20110706053514/http://ods.od.nih.gov/factsheets/VitaminD-HealthProfessional/ |archive-date=6 July 2011 |website=[[Office of Dietary Supplements]], [[US National Institutes of Health]] |url-status=live |access-date=17 August 2021}}</ref> Undernutrition can lead to [[wasting]] in acute cases, and [[stunted growth|stunting]] of [[marasmus]] in chronic cases of malnutrition.<ref name="Whitney-Rolfes-lead" />

==== Domesticated animal ====
In [[domesticated animal]]s, such as [[pet]]s, [[livestock]], and [[working animal]]s, as well as other [[Captivity (animal)|animals in captivity]], nutrition is managed by humans through [[animal feed]]. [[Fodder]] and [[forage]] are provided to livestock. Specialized [[pet food]] has been manufactured since 1860, and subsequent research and development have addressed the nutritional needs of pets. [[Dog food]] and [[cat food]] in particular are heavily studied and typically include all essential nutrients for these animals. Cats are sensitive to some common nutrients, such as [[taurine]], and require additional nutrients derived from meat. Large-breed puppies are susceptible to overnutrition, as small-breed dog food is more energy dense than they can absorb.<ref>{{Cite book |last=Taylor |first=Mark B. |title=Caring for Family Pets: Choosing and Keeping Our Companion Animals Healthy |publisher=[[ABC-CLIO]] |year=2011 |isbn=978-0-313-38527-8 |editor-last=Davis |editor-first=Radford G. |pages=177–194 |chapter=Pet Nutrition}}</ref>

=== Plant ===
{{Main|Plant nutrition}}
[[File:Photosynthesis en.svg|thumb|Schematic of photosynthesis in plants. The carbohydrates produced are stored in or used by the plant.]]
Most plants obtain nutrients through inorganic substances [[Plant nutrients in soil|absorbed from the soil]] or the atmosphere. Carbon, hydrogen, oxygen, nitrogen, and sulfur are essential nutrients that make up organic material in a plant and allow enzymic processes. These are absorbed ions in the soil, such as [[bicarbonate]], [[nitrate]], [[ammonium]], and [[sulfate]], or they are absorbed as gases, such as carbon dioxide, water, oxygen gas, and [[sulfur dioxide]]. Phosphorus, boron, and silicon are used for [[esterification]]. They are obtained through the soil as [[phosphate]]s, [[boric acid]], and [[silicic acid]], respectively. Other nutrients used by plants are potassium, sodium, calcium, magnesium, manganese, chlorine, iron, copper, zinc, and molybdenum.{{Sfn|Mengel|Kirkby|Kosegarten|Appel|2001|pp=1–3}}

Plants uptake essential elements from the soil through their [[root]]s and from the air (consisting of mainly nitrogen and oxygen) through their [[Leaf|leaves]]. Nutrient uptake in the soil is achieved by [[cation exchange]], wherein [[root hair]]s pump [[hydrogen ion]]s (H<sup>+</sup>) into the soil through [[proton pump]]s. These hydrogen ions displace [[cation]]s attached to negatively charged soil particles so that the cations are available for uptake by the root. In the leaves, [[stomata]] open to take in carbon dioxide and expel [[oxygen]].{{Sfn|Mengel|Kirkby|Kosegarten|Appel|2001|pp=111–135}} Although [[nitrogen]] is plentiful in the Earth's atmosphere, very few plants can use this directly. Most plants, therefore, require nitrogen compounds to be present in the soil in which they grow. This is made possible by the fact that largely inert atmospheric nitrogen is changed in a [[nitrogen fixation]] process to biologically usable forms in the soil by bacteria.<ref>{{cite web |last1=Lindemann |first1=W.C. |last2=Glover |first2=C.R. |date=2003 |url=http://aces.nmsu.edu/pubs/_a/A129/welcome.html |title=Nitrogen Fixation by Legumes |archive-url=https://web.archive.org/web/20130517232720/http://aces.nmsu.edu/pubs/_a/A129/welcome.html |archive-date=17 May 2013 |website=[[New Mexico State University]]}}</ref>

As these nutrients do not provide the plant with energy, they must obtain energy by other means. [[Green plant]]s absorb energy from [[sunlight]] with [[chloroplasts]] and convert it to usable energy through [[photosynthesis]].{{Sfn|Mengel|Kirkby|Kosegarten|Appel|2001|pp=136–137}}

=== Fungus ===
{{See also|Fungiculture}}
Fungi are chemoheterotrophs that consume external matter for energy. Most fungi absorb matter through the root-like mycelium, which grows through the organism's source of nutrients and can extend indefinitely. The fungus excretes [[Fungal extracellular enzyme activity|extracellular enzymes]] to break down surrounding matter and then absorbs the nutrients through the cell wall. Fungi can be parasitic, saprophytic, or symbiotic. Parasitic fungi attach and feed on living hosts, such as animals, plants, or other fungi. Saprophytic fungi feed on dead and decomposing organisms. Symbiotic fungi grow around other organisms and exchange nutrients with them.<ref>{{Cite book |last=Charya |first=M. A. Singara |title=Plant Biology and Biotechnology |publisher=[[Springer Science+Business Media|Springer]] |year=2015 |isbn=978-81-322-2286-6 |editor1-last=Bahadur |editor1-first=Bir |pages=197–215 |chapter=Fungi: An Overview |doi=10.1007/978-81-322-2286-6_7 |editor2-last=Rajam |editor2-first=Manchikatla Venkat |editor3-last=Sahijram |editor3-first=Leela |editor4-last=Krishnamurthy |editor4-first=K. V.}}</ref>

=== Protist ===
[[Protist]]s include all [[eukaryote]]s that are not animals, plants, or fungi, resulting in great diversity between them. [[Algae]] are photosynthetic protists that can produce energy from light. Several types of protists use mycelium similar to those of fungi. [[Protozoa]] are heterotrophic protists, and different protozoa seek nutrients in different ways. [[Flagellate]] protozoa use a [[flagellum]] to assist in hunting for food, and some protozoa travel via infectious spores to act as parasites.<ref>{{Cite book |title=Handbook of the Protists |publisher=[[Springer Science+Business Media|Springer]] |year=2017 |isbn=978-3-319-28149-0 |editor1-last=Archibald |editor1-first=John M. |edition=2nd |pages=2–4 |doi=10.1007/978-3-319-28149-0 |lccn=2017945328 |editor2-last=Simpson |editor2-first=Alastair G. B. |editor3-last=Slamovits |editor3-first=Claudio H.}}</ref> Many protists are mixotrophic, having both phototrophic and heterotrophic characteristics. Mixotrophic protists will typically depend on one source of nutrients while using the other as a supplemental source or a temporary alternative when its primary source is unavailable.<ref>{{Cite journal |last=Jones |first=Harriet |date=1997 |title=A classification of mixotrophic protists based on their behaviour |journal=Freshwater Biology |volume=37 |issue=1 |pages=35–43 |doi=10.1046/j.1365-2427.1997.00138.x |bibcode=1997FrBio..37...35J }}</ref>

=== Prokaryote ===
{{Further|Microbial metabolism}}
[[File:Metabolism-en.svg|thumb|Simplified view of [[Metabolism|cellular metabolism]]]]
[[Prokaryote]]s, including [[bacteria]] and [[archaea]], vary greatly in how they obtain nutrients across nutritional groups. Prokaryotes can only transport soluble compounds across their cell envelopes, but they can break down chemical components around them. Some lithotrophic prokaryotes are [[extremophile]]s that can survive in nutrient-deprived environments by breaking down inorganic matter.<ref>{{cite book |doi=10.1016/B978-044452748-6.00164-4 |chapter=Geology, Life and Habitability |title=Treatise on Geophysics |date=2007 |last1=Southam |first1=G. |last2=Westall |first2=F. |pages=421–437 |isbn=978-0-444-52748-6 |chapter-url=https://elib.dlr.de/97967/1/10-14-Geology-Life-and-Habitability_2015_Treatise-on-Geophysics-Second-Edition-.pdf }}</ref> Phototrophic prokaryotes, such as [[cyanobacteria]] and [[Chloroflexia]], can engage in photosynthesis to obtain energy from sunlight. This is common among bacteria that form in mats atop geothermal springs. Phototrophic prokaryotes typically obtain carbon from assimilating carbon dioxide through the [[Calvin cycle]].<ref>{{cite book |doi=10.1007/0-387-30742-7_3 |chapter=The Phototrophic Way of Life |title=The Prokaryotes |date=2006 |last1=Overmann |first1=Jörg |last2=Garcia-Pichel |first2=Ferrau |pages=32–85 |isbn=978-0-387-25492-0 }}</ref>

Some prokaryotes, such as ''[[Bdellovibrio]]'' and [[Ensifer (bacterium)|''Ensifer'']], are predatory and feed on other single-celled organisms. Predatory prokaryotes seek out other organisms through [[chemotaxis]] or random collision, merge with the organism, degrade it, and absorb the released nutrients. Predatory strategies of prokaryotes include attaching to the outer surface of the organism and degrading it externally, entering the cytoplasm of the organism, or by entering the [[Periplasm|periplasmic space]] of the organism. Groups of predatory prokaryotes may forgo attachment by collectively producing [[Hydrolysis|hydrolytic]] enzymes.<ref>{{cite journal |last1=Martin |first1=Mark O. |title=Predatory prokaryotes: an emerging research opportunity |journal=Journal of Molecular Microbiology and Biotechnology |date=September 2002 |volume=4 |issue=5 |pages=467–477 |pmid=12432957 }}</ref>

==See also==
* {{annotated link|Liebig's law of the minimum}}
* {{annotated link|Nutrient density}}
* {{annotated link|Nutrition analysis}}
* {{annotated link|Resource (biology)}}
* {{annotated link|Substrate (biology)}}
* [[Milan Charter]] 2015 Charter on Nutrition

== References ==
{{Reflist}}

== Bibliography ==
* {{Cite book |last=Andrews |first=John H. |title=Comparative Ecology of Microorganisms and Macroorganisms |publisher=[[Springer Science+Business Media|Springer]] |location=New York |year=2017 |isbn=978-1-4939-6897-8 |edition=2nd}}
* {{cite book |last1=Callahan |first1=Alice |last2=Leonard |first2=Heather |last3=Powell |first3=Tamberly |title=Nutrition: Science and Everyday Application |url=https://openoregon.pressbooks.pub/nutritionscience2e/ |year=2022 |publisher=[[Pressbooks]]}}
* {{Cite book |editor1-last=Mann |editor1-first=Jim |title=Essentials of Human Nutrition |publisher=[[Oxford University Press]] |location=Oxford |year=2012 |isbn=978-0-19-956634-1 |edition=4th |editor2-last=Truswell |editor2-first=A. Stewart}}
* {{Cite book |editor1-last=Mengel |editor1-first=Konrad |title=Principles of Plant Nutrition |publisher=[[Springer Science+Business Media|Springer]] |location=New York |year=2001 |isbn=978-94-010-1009-2 |edition=5th |doi=10.1007/978-94-010-1009-2 |editor2-last=Kirkby |editor2-first=Ernest A. |editor3-last=Kosegarten |editor3-first=Harald |editor4-last=Appel |editor4-first=Thomas}}
* {{Cite book |last1=Simpson |first1=Stephen J. |title=The Nature of Nutrition: A Unifying Framework from Animal Adaptation to Human Obesity |last2=Raubenheimer |first2=David |publisher=[[Princeton University Press]] |location=Princeton |year=2012 |isbn=978-1-4008-4280-3}}
* {{Cite book |last=Wu |first=Guoyao |title=Principles of Animal Nutrition |publisher=[[CRC Press]] |location=Boca Raton |year=2017 |isbn=978-1-351-64637-6}}

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